比利（丹尼斯·霍珀 Dennis Hopper 饰）和怀特（彼得·方达 Peter Fonda 饰）用一次毒品交易的钱骑着他们的机车上路了，他们的目的地，是新奥尔良的狂欢节。在路上，他们经过了离群索居的波西米亚人群落，短暂逗留的数天里目睹了他们自由放浪但也贫穷困苦的生活。在德克萨斯州，他们仅因为行为举止诡异就被警察关进了监狱，在那里，他们遇见了律师汉森（杰克·尼科尔森 Jack Nicholson 饰），汉森帮助他们逃离了牢狱之灾，他们怂恿汉森同他们一起上路。抵挡不住内心的骚动，汉森同意了。三人来到一个保守的小镇，在那里受尽了居民的白眼。不受欢迎的三人只得露宿荒野，夜里，当地的居民乔装袭击了他们，汉森被打死。比利和怀特虽然震惊，但这一切都没能阻止他们前进的脚步。到达目的地的两人并没有感到喜悦，有的只是无尽的空虚，沮丧的两人打道回府，但令人意外的是，一个卡车司机和他的双杆猎枪使得他们的行程截然而止。 这是影史上最伟大的公路电影，也是为数众多的公路电影的鼻祖。
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